NY市警での一件から数日… 5年前の惨劇を止められなかったことを悔やむレオに ガンビーノ=ファミリーの家長、ヨハン=ガンビーノが接触する。 ヨハンは腐りきったこの社会をひっくり返すべく、 絶大な力を持つレオの「グランファミリア」の指輪を狙い抗争を仕掛ける! 特殊な能力を使い人々を混乱に陥れるガンビーノ=ファミリー! 亜人ファミリー同士の抗争が始まる!
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